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Goffinet, B.; Shaw, A.J.; Cox, C.J.; Wickett, N.J.; Boles, S.B. 2004: Phylogenetic inferences in the Orthotrichoideae (Orthotrichaceae, Bryophyta) based on variation in four loci from all genomes. In: Hollowell, V.B (ed.) Molecular Systematics of Bryophytes. Vol. 98. In: Monographs in Systematic Botany from the Missouri Botanical Garden. 270–289.